DAVID E CADE
  • Home
  • Research
    • Field Projects
  • Publications
  • Teaching
  • Outreach
    • Media
  • Collaborations
  • CV
Allen, J.A., Cade, D.E., Casey, C.E., Weindor, S., Linsky, J.M.J., Goldbogen, J.A., Nowacek, D.P., & Friedlaender, A.S (2024). Evidence of sociality and group foraging in Antarctic minke whales (Balaenoptera bonaerensis). Behavioral Ecology and Sociobiology, 78(5), 1-14.

Bamford, C.C.G., Kelly, N., Dalla Rosa, L., Cade, D.E., Fretwell, P., Tratham, P.N., Cubaynes, H., Mesquita, A., Gerrish, L., Friedlaender, A.S., Jackson, J.A. (2020) A comparison of baleen whale density estimates derived from overlapping satellite imagery and a shipborne survey. Scientific Reports, 10, 1-12.  

Blawas, A.M., Videsenn, S.K.A., Cade, D.E., Calambokidis, J., Friedlaender, A.S., Johnston, D.W., Madsen, P.T., Goldbogen, J.A. (2025). Life in the slowest lane: Feeding allometry lowers metabolic rate scaling in the largest whales. Science Advances, 11(32), eadw2232.

Cade, D.E., Barr, K.R., Calambokidis, J., Friedlaender, A.S. & Goldbogen, J.A. (2018) Determining forward speed from accelerometer jiggle in aquatic environments. Journal of Experimental Biology, 221, jeb170449.

Cade, D.E. & Benoit-Bird, K.J. (2014) An automatic and quantitative approach to the detection and tracking of acoustic scattering layers. Limnology and Oceanography: Methods, 12, 742-756.

Cade, D.E. & Benoit-Bird, K.J. (2015) Depths, migration rates and environmental associations of acoustic scattering layers in the Gulf of California. Deep Sea Research Part I: Oceanographic Research Papers, 102, 78-89.

Cade, D.E., Carey, N., Domenici, P., Potvin, J. & Goldbogen, J.A. (2020a) Predator-informed looming stimulus experiments reveal how large filter feeding whales capture highly maneuverable forage fish. Proceedings of the National Academy of Sciences, 117, 472-478.

Cade, D.E., Fahlbusch, J.A., Oestreich, W.K., Ryan, J., Calambokidis, J., FIndlay, K.P, Friedlaender, A.S., Hazen, E.S., Seakamela, S.M. & Goldbogen, J.A. (2021) Social exploitation of extensive, ephemeral, environmentally controlled prey patches by super groups of rorqul whales. Animal Behaviour, 182, 251-266.

Cade, D.E., Friedlaender, A.S., Calambokidis, J. & Goldbogen, J.A. (2016) Kinematic Diversity in Rorqual Whale Feeding Mechanisms. Current Biology, 26, 2617-2624.

Cade, D.E., Gough, W.T., Czapanskiy, M.F., Fahlbusch, J.A., Kahane-Rapport, S.R., Linsky, J.M.J., Nichols, R.C., Oestreich, W.K., Wisniewska, D.M., Friedlaender, A.S., & Goldbogen, J.A. (2021) Tools for integrating inertial sensor data with video bio-loggers, including estimation of animal orientation, motion and position.  Animal Biotelemetry, 9, 34.

Cade, D.E., Kahane-Rapport, S.R., Gough, W.T., Bierlich, K.C., Linsky, J.M.L., Calambolidis, J., Johnston, D.W., Goldbogen, J.A. & Friedlaender, A.S. (2023) Minke whale feeding rate limitations suggest constraints on the minimum body size for engulfment filtration feeding. Nature: Ecology & Evolution, 7, 535-546.

Cade, D.E., Kahane-Rapport, S.R., Wallis, B., Goldbogen, J.A. & Friedlaender, A.S. (2022) Evidence for size-selective predation in Antarctic humpback whales.  Frontiers in Marine Science, 9, 747788.

Cade, D.E., Levenson, J.J., Cooper, B., de la Parra, R., Webb, D.H., & Dove, A. (2020b) Whale sharks increase swimming effort while filter feeding, but appear to maintain high foraging efficiencies. Journal of Experimental Biology, 223, jeb224402.

Cade, D.E., Seakamela, S.M., Findlay, K,P., Fukunaga, J., Kahane-Rapport, S.R., Warren, J.D., Calambokidis, J., Fahlbusch, J.A., Friedlaender, A.S., Hazen, E., Kotze, D., McCue, S., Meyer, M., Oestreich, W.K., Oudejans, M., Wilke, C. & Goldbogen, J.A. (2021) Predator scale spatial analysis of intra-patch prey distribution reveals the energetic drivers of rorqual whale super-group formation. Functional Ecology, 35, 894-908


Calambokidis, J., Fahlbusch, J.A., Szesciorka, A.R., Southall, B.L., Cade, D.E., Friedlaender, A.S. & Goldbogen, J.A. (2019) Differential Vulnerability to Ship Strikes between Day and Night for Blue, Fin, and Humpback Whales Based on Dive and Movement Data from Medium Duration Archival Tags. Frontiers in Marine Science, 6.

Casey, C.B., Weindorf, S., Levy, E., Linsky, J.M.J., Cade, D.E., Goldbogen, J.A., Nowacek, D.P., Friedlaender, A.S. (2022) Acoustic signalling and behavior of Antarctic minke whales (Balaenoptera bonaerensis). Royal society open science, 9, 211557.

Chapple, T., Cade, D.E., Goldbogen, J.A., Massett, N., Payne, N.L., McInturf, A.G. (2024). Behavioral response of megafauna to boat collision measured via animal-borne camera and IMU. Frontiers in Marine Science, 11, 1430961.

Clayton, H., Cade, D.E., Burnham, R., Calambokidis, J. Goldbogen, J.A. (2023) Acoustic behavior of gray whales tagged with biologging devices on foraging grounds. Frontiers in Marine Science, 10, 1111666.

Colson, K,M., Pirotta, E., New, L., Cade, D.E., Calambokidis, J., Bierlich, K.C., Bird, C.N., Ajo, A.F., Hildebrand, L., Trites, A.W., Torres, L.G. (2024). Using accelerometry tags to quantify gray whale foraging behavior. Marine Mammal Science. e13210.

Czapanskiy, M.F., Savoca, M.S., Gough, W.T., Segre, P.S., Wisniewska, D.M., Cade, D.E., & Goldbogen, J.A. (2019) Modeling short-term energetic costs of sonar disturbance to cetaceans using high-resolution foraging data. Journal of Applied Ecology, 58, 1643-1657

Fahlbusch, J.A., Czapanskiy, M.F., Calambokidis, J., Cade, D.E., Abrahms, B., Hazen, E.L., Goldbogen, J.A. (2022) Blue whales increase feeding rates at fine-scale ocean features. Proceedings of the Royal Society B: Biological Sciences,289, 20221180.


Fahlbusch, J.A., Cade, D.E., Haxen, E., Elliott, M., Saenz, B., Goldbogen, J.A., Jahncke, J. (2024). Submesoscale physical-biological coupling between krill and cetaceans in the CCS. Proceedings of the Royal Society B: Biological Sciences, 291(2017), 20232461

Findlay, K.P., Seakamela, S.M., Meÿer, M.A., Kirkman, S.P., Barendse, J., Cade, D.E., Hurwitz, D., Kennedy, A., Kotze, P.G.H., McCue, S.A., Thornton, M., Vargas-Fonseca, O.A. & Wilke, C.G. (2017) Humpback whale “super-groups” – A novel low-latitude feeding behaviour of Southern Hemisphere humpback whales (Megaptera novaeangliae) in the Benguela Upwelling System. PloS one, 12, e0172002.

Flammang, B.E., Marras, S., Anderson, E.J., Lehmkuhl, O., Mukherjee, A., Cade, D.E., Beckert, M., Nadler, J.H., Houzeaux, G., Vazquez, M., Amplo, H.E., Calambokidis, J, Friedlaender, A.S., Goldbogen, J.A. (2020) Remoras pick where they stick on blue whales. Journal of Experimental Biology, 223, 1-12.

Friedlaender, A.S., Bowers, M.T., Cade, D., Hazen, E.L., Stimpert, A.K., Allen, A.N., Calambokidis, J., Fahlbusch, J., Segre, P. & Visser, F. (2020) The advantages of diving deep: Fin whales quadruple their energy intake when targeting deep krill patches. Functional Ecology.


Friedlaender, A.S., Herbert-Read, J.E., Hazen, E.L., Cade, D.E., Calambokidis, J., Southall, B.L., Stimpert, A.K. & Goldbogen, J.A. (2017) Context-dependent lateralized feeding strategies in blue whales. Current Biology, 27, R1206-R1208.

Goldbogen, J.A., Cade, D.E., (2025). How do feeding biomechanics, extreme predator-prey size ratios, and the rare enemy effect determine energetics at the largest scale? Journal of Experimental Biology, jeb247875.

Goldbogen, J.A., Cade, D.E., Boersma, A.T., Calambokidis, J., Kahane‐Rapport, S.R., Segre, P.S., Stimpert, A.K. & Friedlaender, A.S. (2017a) Using Digital Tags With Integrated Video and Inertial Sensors to Study Moving Morphology and Associated Function in Large Aquatic Vertebrates. The Anatomical Record, 300, 1935-1941.

Goldbogen, J.A., Cade, D.E., Calambokidis, J., Czapanskiy, M.F., Fahlbusch, J., Friedlaender, A.S., Gough, W.T., Kahane-Rapport, S.R., Savoca, M.S. & Ponganis, K.V. (2019a) Extreme bradycardia and tachycardia in the world’s largest animal. Proceedings of the National Academy of Sciences, 116, 25329-25332.

Goldbogen, J.A., Cade, D.E., Calambokidis, J., Friedlaender, A.S., Potvin, J., Segre, P.S. & Werth, A.J. (2017b) How Baleen Whales Feed: The Biomechanics of Engulfment and Filtration. Annual review of marine science, 9, 1-20.

Goldbogen, J.A., Cade, D.E., Wisniewska, D.M., Potvin, J., Segre, P., Savoca, M.S., Hazen, E.L., Czapanskiy, M.F., Kahane-Rapport, S.R., DeRuiter, S.L., Gero, S., Tønnesen, P., Gough, W.T., Hanson, M.B., Holt, M., Jensen, F.H., Simon, M., Stimpert, A.K., Arranz, P., Nowacek, D.P., Parks, S., Visser, F., Friedlaender, A.F., Tyack, P.L., Madsen, P.T. & Pyenson, N.D. (2019b) Why whales are big but not bigger: physiological drivers and ecological limits in the age of ocean giants. Science, 366, 1367-1372.

Gough, W.T., Segre, P.S., Bierlich, K., Cade, D.E., Potvin, J., Fish, F.E., Dale, J., di Clemente, J., Friedlaender, A.S., Johnston, D.W., Kahane-Rapport, S., Kennedy, J., Long, J., Oudejans, M., Penry, G.S., Savoca, M.S., Simon, M., Videsen, S., Visser, F., Wiley, D., & Goldbogen, J.A.. (2019) Scaling of swimming performance in baleen whales. Journal of Experimental Biology, 222, jeb. 204172.

Gough, W.T., Smith, H.J., Savoca, M.S., Czapanskiy, M.F., Fish, F.E., Potvin, J., Bierlich, K.C., Cade, D.E., di Clemente, J., Kennedy, J., Segre, P.S., Stanworth, A., Weir, C., & Goldbogen, J.A. (2021) Scaling of oscillatory kinematics and Froude efficiency in baleen whales. Journal of Experimental Biology, 224, jeb. 237586.

Gough, W.T., Cade, D.E., Czapanskiy, M.F., Potvin, J., Fish, Frank E., Kahane-Rapport, S.R., Savoca, M.S., Bierlich, K.C., Johnston, D.W., et al. (2022) Fast and furious: Energetic tradeoffs and scaling of high-speed foraging in rorqual whales. Integrative and Organismal Biology, 4, jeb. obac038.

Hein, A.M., Altshuler, D. L., Cade, D.E., Liao, J.C., Martin, B.T., Taylor, G.K. (2020) An algorithmic approach to natural behavior.  Current Biology, 30, R663-R675. 

Kahane-Rapport, S.R., Savoca, M.S., Cade, D.E., Segre, P.S., Bierlich, K.C., Calambokidis, J., Dale, J., Fahlbusch, J.A., Friedlaender, A.S., Johnston, D., Werth, A.J.,. & Goldbogen, J.A. (2020) Lunge filter-feeding biomechanics constrain rorqual foraging ecology across scale. Journal of Experimental Biology, jeb. 224196.


Li, T., Schindler, M., Paskin, M., Surapaneni, V., Scott, E., Hauert, S., Payne, N., Cade, D.E., Goldbogen, J.A., Mollen, F.H., Baum, D., Hanna, S., Dean, M.N. (2025). Functional Models, from limited data: a parametric and multimodal approach to anatomy and 3D kinematics of feeding in basking sharks (Cetorhinus maximus). The Anatomical Record. DOI: 10.1002/ar.25693.


Linsky, J., Wilson, N, Cade, D.E., Goldbogen, J.A., Johnston, D.W., Friedlaender, A.S. (2020) The scale of the whale: using video-tag data to evaluate sea-surface ice concentration from the perspective of individual Antarctic minke whales. Animal Biotelemetry, 8, 1-12.

Martin-Lopez, L.M., Isojunno, S., Cade, D.E., Colson, K., Paradinas, I., Miller, P.J.O., Fahlman, A., Hickmott, L.S., Visser, F., (2025). Naval sonar induces an anaerobic swimming gait in beaked whales. Scientific Reports, 15 (38686).

Mastick, N., Wiley, D., Cade, D.E., Ware, C., Parks, S.E., Friedlaender, A.S. (2022) The effect of group size on individual behaviors of bubble-net feeding humpback whales in the southern Gulf of Maine. Marine Mammal Science, 38, 959-974.


Matika, A.F., Jourdain, E., Cade, D.E., Karoliussen, R., Hammond, P.S. (2022) Feeding characteristics and prey profitability in five herring-feeding killer whales (Orcinus Orca) in Northern Norway. Marine Mammal Science, 38, 1409-1424.

Nazario, E., Cade, D.E., Bierlich, K.C., Czapanskiy, M.F., Goldbogen, J.A., Kahane-Rapport, S.R., van der Hoop, J.M., San Luis, M.T., Friedlaender, A.S. (2022) Baleen whale inhalation variability revealed using animal-borne video tags. PeerJ, 10, 13724.

Nichols, R.C., Cade, D.E., Kahane-Rapport, S.R., Goldbogen, J.A., Stimpert, A.S., Nowacek, D., Read., A.J., Johnston, D.W., Friedlaender, A.S. (2022) Intra-seasonal variation in feeding-rates and diel foraging behaviour of a seasonally fasting mammal, the humpback whale. Royal Society Open Science, 9, 211674.

Oestrich, W. K. Fahlbusch, J. A., Cade, D. E. , Calambokidis, J., Margolina, T., Joseph, J., Friedlaender, A.S., McKenna, M.F., Stimpert, A.K., Southall, B.L., Goldbogen, J.A., Ryan, J.P. (2020) Animal-borne metrics enable acoustic detection of blue whale mibgration. Current Biology, 30, 1-7.

Payne, N.L., Snelling, E.P., Peralta-Maraver, I., Cade, D.E., Chapple, T.K., McInturf, A.G., Watanabe, Y.Y., Sims, D.W., Queiroz, N., da Costa, I., Sousa, L.L, Goldbogen, J.A., Dolton, H.R., Jackson, A.L. (2026). Mesothermic fishes face high fuel demands and overheating risks in warming oceans. Science, 392(6795), 301-305.

Potvin, J. Cade, D.E., Werth, A.J., Shadwick, R.E., Goldbogen, J.A. (2020) A perfectly inelastic collision: Bulk prey engulfment by baleen whales and dybamical implications for the world's largest cetaceans. American Journal of Physics, 88, 851-863. 

Potvin, J. Cade, D.E., Werth, A.J., Shadwick, R.E., Goldbogen, J.A. (2021) Rorqual lunge-feeding energetics near and away from the kinematic threshold of optimal efficiency. Integrative and Organismal Biology, 3, obab005. 


Price, S.E., Savoca, M.S., Kumar, M., Czapanskiy, M.F., McDermott, D., Litvin, S.Y., Cade, D.E., Goldbogen, J.A. (2024). Energy Densities of key prey species in the California Current Ecosystem. Frontiers in Marine Science. 10, 1345525.

Ryan, J., Benoit-Bird, K.J., Oestrich, W.K., Leary, P., Smith, K., Waluk, C.P., Cade, D.E., Fahlbusch, J.A., Southall, B., et al. (2022) Oceanic giants dance to atmospheric rhythms: Ephemeral wind-driven resource tracking by blue whales. Ecology Letters, 5 Oct 2022, 1-13.


Segre, P.S., Cade, D.E., Calambokidis, J., Fish, F.E., Friedlaender, A.S., Potvin, J. & Goldbogen, J.A. (2018) Body flexibility enhances maneuverability in the world’s largest predator. Integrative and comparative biology. 59, 48-60.

Segre, P.S., Cade, D.E., Fish, F.E., Potvin, J., Allen, A.N., Calambokidis, J., Friedlaender, A.S. & Goldbogen, J.A. (2016) Hydrodynamic properties of fin whale flippers predict maximum rolling performance. Journal of Experimental Biology, 219, 3315-3320.


Segre, P.S., Gough, W.T., Roualdes, E.A., Cade, D.E., Czapanskiy, M.F., et al., (2022) Scaling of maneuvering performance in baleen whales: larger whales outperform expectations. Journal of Experimental Biology, 225, jeb243224.

Segre, P.S., Potvin, J., Cade, D.E., Calambokidis, J., Di Clemente, J., Fish, F.E., Friedlaender, A.S., Gough, W.T., Kahane-Rapport, S.R. & Oliveira, C. (2020) Energetic and physical limitations on the breaching performance of large whales. eLife, 9.

Southall, B.L., DeRuiter, S.L., Friedlaender, A., Stimpert, A.K., Goldbogen, J.A., Hazen, E., Casey, C., Fregosi, S., Cade, D.E. & Allen, A.N. (2019) Behavioral responses of individual blue whales (Balaenoptera musculus) to mid-frequency military sonar. Journal of Experimental Biology, 222, jeb190637.

Tackaberry, J.E., Cade, D.E., Goldbogen, J.A., Wiley, D.N., Friedlaender, A.S., Stimpert, A.S. (2020) From a calf's perspective: Humpback whale nursing behavior on two US feeding grounds.  Peer J, 
8:e8538.

Werth, A.J., Potvin, J., Shadwick, R.E., Jensen, M.M., Cade, D.E. & Goldbogen, J.A. (2018) Filtration area scaling and evolution in mysticetes: trophic niche partitioning and the curious cases of sei and pygmy right whales. Biological Journal of the Linnean Society, 125, 264-279
 

Cover
articles

Picture
Cade et al. 2023. Minke whale feeding rate limitations suggest constraints on the minimum body size for engulfment filtration feeding

Picture
Cade et al. 2020. Predator-informed looming stimulus experiments reveal how large filter feeding whales capture highly maneuverable forage fish.
A discussion with Dr. Cade
Picture

Picture
Cade et al. 2020. Whale sharks increase swimming effort while filter feeding, but appear to maintain high foraging efficiencies.
Resumen en español

Picture
Ryan et al., 2022. Blue whale vocalizations track with ocean processes

Picture
Segre et al. 2022. Scaling of maneuvering performance in baleen whales: large whales outperform expectations

Picture
Mastick et al. 2022. Effects of group size on bubble-net feeding

Picture
Cade et al. 2016. Kinematic diversity in rorqual whale feeding mechanisms

Picture
Goldbogen, Cade et al. 2019. Why whales are big but not bigger: Physiological drivers and ecological limits in the age of ocean giants

Picture
Friedlaender et al. 2017. Context-dependent lateralized feeding strategies in blue whales

Other Media

Picture
Goldbogen, Cade et al. (2019) PNAS. First report of the extreme bradycardia and tachycardia in the world's largest animal
(above) discussion with Dr. Cade on the CBC's Quirks and Quarks

Supplemental videos for Cade, Carey et al. (2020) PNAS

Picture
Part I- Humpback whales in the study fed at high speeds on small schools in concert with common dolphins. Part II- A humpback whale from Monterey Bay, California fed at slower speeds on a large school in concert with California sea lions.
Picture
Part I- Humpback whale flippers are extraordinarily long (~ 30% of body length) with white bottoms, and attacking whales regularly rotate and extend them when lunge-feeding, a technique that has been shown to have a startle effect on prey. Part II- When humpback whales approach prey but do not open their mouths, schools maintain formation since the threshold of response is not reached
Picture
An anchovy demonstrates a characteristic C-start escape response away from the stimulus when startled. High-speed cameras (250 Hz) allowed precise determination of the timing of response, and the frame number in the corner of the playback screen showed the stage of the animation at which the response occurred.

Supplemental videos for
Cade et al. (2018) J. Exp Bio

Supplemental videos for
Cade et al. (2016) Current Biology

Proudly powered by Weebly
  • Home
  • Research
    • Field Projects
  • Publications
  • Teaching
  • Outreach
    • Media
  • Collaborations
  • CV